Coossification

Given the relative size of the specimens, the level of coossification between the quadrate and quadratojugal and the development of dorsal and mandibular condyles, it is assumed that Morphotype 1 includes quadrates belonging to juvenile, immature, sub-adult and adult individuals, so that a sequence list of ontogenetic character transformations (maturity-dependent characters) can be provided for the quadrate of Spinosaurus aegyptiacus.