Holaspis

A similar situation may also characterize S. wenlockiana, a species with six segments in holaspis, the smallest holaspids of which have similar glabellar lengths to those of the sixth degree meraspis of A. konincki.

The pygidial spines are stout and direct outward in the early holaspis (Fig. 2, A-B, F), but become slender and direct inward in the late holaspis (Fig. 2, D, E).

The holaspis stage has a full complement of thoracic segments for the species but growth continues through further molts and maturity may not be reached until some time after the holaspis stage was reached.