The model itself operates at three time scales: i) a within patch foraging scale (one tick of the intragenerational clock); ii) an intragenerational patch-to-patch movement scale (n ticks of the intrageneration clock equal to 1 tick of the intergenerational clock); and iii) an intergenerational evolutionary scale (G ticks of the epochal clock), as depicted in Fig 2 .
Source: wiktionary