3) Stages of radiation: All else being equal, the following sequence of the diversification events is expected: (i) divergence with respect to macrohabitat; (ii) evolution of microhabitat choice and divergence with respect to microhabitat; (iii) divergence with respect to traits that simultaneously control the degree of local adaptation and nonrandom mating; and (iv) divergence with respect to other traits controlling survival and reproduction.
Source: wiktionary